Graduated in geology (BSc) at the University of Tartu in 1993. MSc in paleontology and stratigraphy from the University of Tartu in 1995. PhD in geology from the University of Tartu in 2001. From 2001 to 2007 researcher, and since 2007 senior researcher at the Department of Geology, University of Tartu. Main research interests are: (1) tube ultrastructure and evolution of biomineralization in annelid tubeworms (both fossil and recent); (2) biological affinities, shell structure and evolution of problematic tubicolous fossils from Ediacaran and Phanerozoic; (3) bioerosion and biofouling, palaeoecology and evolution during the Phanerozoic; (4) evolution of symbiosis and predation.